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Grizzly Bear (Ursus arctos horribilis) footprint in snow next to human print, central Alaska
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00558003 Grizzly Bear (Ursus arctos horribilis) footprint in snow next to human print, central Alaska© Donald M. Jones
Javan Rhinoceros (Rhinoceros sondaicus) foot print being examined by Rhino Protection Unit Ranger, Ujung Kulon National Park, Indonesia  -  Stephen Belcher
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00542090 Javan Rhinoceros (Rhinoceros sondaicus) foot print being examined by Rhino Protection Unit Ranger, Ujung Kulon National Park, Indonesia© Stephen Belcher
Spanish Lynx (Lynx pardinus) track and human hand print, Donana National Park, Huelva, Andalusia, Spain  -  Pete Oxford
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00442297 Spanish Lynx (Lynx pardinus) track and human hand print, Donana National Park, Huelva, Andalusia, Spain© Pete Oxford
Sumatran Rhinoceros (Dicerorhinus sumatrensis) plaster cast of juvenile and mature sumatran rhino foot print, Way Kambas National Park, Sumatra, Indonesia  -  Stephen Belcher
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00436772 Sumatran Rhinoceros (Dicerorhinus sumatrensis) plaster cast of juvenile and mature sumatran rhino foot print, Way Kambas National Park, Sumatra, Indonesia© Stephen Belcher
Sumatran Rhinoceros (Dicerorhinus sumatrensis) plaster cast of juvenile foot print, Way Kambas National Park, Sumatra, Indonesia  -  Stephen Belcher
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00436770 Sumatran Rhinoceros (Dicerorhinus sumatrensis) plaster cast of juvenile foot print, Way Kambas National Park, Sumatra, Indonesia© Stephen Belcher
Eurasian Kestrel (Falco tinnunculus) print left in snow after catching a mouse, Germany  -  Konrad Wothe
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00429963 Eurasian Kestrel (Falco tinnunculus) print left in snow after catching a mouse, Germany© Konrad Wothe
Orca (Orcinus orca) vocalization print outs from John Ford's research, British Columbia, Canada  -  Flip Nicklin
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00079419 Orca (Orcinus orca) vocalization print outs from John Ford's research, British Columbia, Canada© Flip Nicklin
Long exposure of light trail from bioluminescent click beetle (Pyrophorus sp) walking on ground at night. Danum Valley Sabah, Borneo.
90722570
90722570 Long exposure of light trail from bioluminescent click beetle (Pyrophorus sp) walking on ground at night. Danum Valley Sabah, Borneo.© Alex Hyde
Highland cow next to road above Malham, Yorkshire, winter.
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90720167 Highland cow next to road above Malham, Yorkshire, winter.© Graham Eaton,Graham Eaton
Footprints of Water vole (Arvicola amphibius) or possibly Brown rat (Rattus norvegicus) on muddy river bank, found during survey for signs of Water voles, near Bude, Cornwall, UK, April  -  Nick Upton/ npl
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90673856 Footprints of Water vole (Arvicola amphibius) or possibly Brown rat (Rattus norvegicus) on muddy river bank, found during survey for signs of Water voles, near Bude, Cornwall, UK, April© Nick Upton / NPL
Bobcat (Lynx rufus) adult, running in snow, Montana, U.S.A., january (captive)  -  Paul Sawer/ FLPA
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80125227 Bobcat (Lynx rufus) adult, running in snow, Montana, U.S.A., january (captive)© Paul Sawer / FLPA
Sumatran Rhinoceros (Dicerorhinus sumatrensis) foot print, Sumatran Rhino Sanctuary, Way Kambas National Park, Sumatra, Indonesia  -  Stephen Belcher
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70005029 Sumatran Rhinoceros (Dicerorhinus sumatrensis) foot print, Sumatran Rhino Sanctuary, Way Kambas National Park, Sumatra, Indonesia© Stephen Belcher
Sumatran Rhinoceros (Dicerorhinus sumatrensis) foot print, Sumatran Rhino Sanctuary, Way Kambas National Park, Sumatra, Indonesia  -  Stephen Belcher
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70005027 Sumatran Rhinoceros (Dicerorhinus sumatrensis) foot print, Sumatran Rhino Sanctuary, Way Kambas National Park, Sumatra, Indonesia© Stephen Belcher
Firefly (Lampyridae) trails on beach at night, Palm tree silhouetted against milky way. Oak Beach, Port Douglas, Far North Queensland, Australia. 2016.
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90852729 Firefly (Lampyridae) trails on beach at night, Palm tree silhouetted against milky way. Oak Beach, Port Douglas, Far North Queensland, Australia. 2016.©
Polar guide from expedition cruise ship snow shoeing on Petermann Island, Wilhelm Archipelago, Antarctica. December 2019.
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90852326 Polar guide from expedition cruise ship snow shoeing on Petermann Island, Wilhelm Archipelago, Antarctica. December 2019.©
Freshly printed Buddhist texts drying. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.
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90850815 Freshly printed Buddhist texts drying. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.©
Craftsman with wooden plate for printing Buddhist prayers. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.
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90850814 Craftsman with wooden plate for printing Buddhist prayers. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.©
Craftsman creating wooden plate for printing Buddhist prayers. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.
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90850813 Craftsman creating wooden plate for printing Buddhist prayers. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.©
Craftsman carving wooden plate for printing Buddhist prayers. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.
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90850812 Craftsman carving wooden plate for printing Buddhist prayers. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.©
Craftsman in workshop with wooden plate for printing Buddhist prayers. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.
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90850811 Craftsman in workshop with wooden plate for printing Buddhist prayers. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.©
Freshly printed Buddhist texts drying at Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.
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90850791 Freshly printed Buddhist texts drying at Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.©
Monk with wooden plates used to print Buddhist prayers and texts. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.
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90850790 Monk with wooden plates used to print Buddhist prayers and texts. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.©
Wooden plates used to print Buddhist prayers. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.
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90850789 Wooden plates used to print Buddhist prayers. Palpung Monastery, Kham, Dege County, Garze Tibetan Autonomous Prefecture, Sichuan, China. 2016.©
Buzzer midge (Chironomus plumosus) swarm at night, movement trails, long exposure. The Netherlands. July.
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90849767 Buzzer midge (Chironomus plumosus) swarm at night, movement trails, long exposure. The Netherlands. July.©
Landscape scarred by peat extraction, aerial view. Skarnes, Innlandet, Norway. August 2019.
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90849355 Landscape scarred by peat extraction, aerial view. Skarnes, Innlandet, Norway. August 2019.©
Island of woodland within field, an example of habitat fragmentation. Ski, Akershus, Viken, Norway. August 2019.
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90849338 Island of woodland within field, an example of habitat fragmentation. Ski, Akershus, Viken, Norway. August 2019.©
Boat and water skier in channel, aerial view. Lofoten, Norway. July 2018.
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90849337 Boat and water skier in channel, aerial view. Lofoten, Norway. July 2018.©
Tyre tracks on sand dunes of Erg Chebbi, shadows in morning light. South Morocco. October 2019.
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90849252 Tyre tracks on sand dunes of Erg Chebbi, shadows in morning light. South Morocco. October 2019.©
Hind paw print of Kamchatka brown bear (Ursus actos) in sand, Bukhta Natalia, Koryaksky Nature Reserve, Kamchatka Peninsula, Russia.
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90842742 Hind paw print of Kamchatka brown bear (Ursus actos) in sand, Bukhta Natalia, Koryaksky Nature Reserve, Kamchatka Peninsula, Russia.© Jenny E. Ross
Tyre tracks in cracked mud of temporary river, Castildetierra mountain rock formation in background, in evening light. Bardenas Reales Natural Park. Navarre. Spain. January 2006
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90840392 Tyre tracks in cracked mud of temporary river, Castildetierra mountain rock formation in background, in evening light. Bardenas Reales Natural Park. Navarre. Spain. January 2006© Eduardo Blanco
African lion (Panthera leo) pride watched by tourists in safari vehicles as viewed from hot air balloon, Masai Mara National Park, Kenya, January.
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90838114 African lion (Panthera leo) pride watched by tourists in safari vehicles as viewed from hot air balloon, Masai Mara National Park, Kenya, January.© Will Watson
A female Humpback whale (Megaptera novaeangliae australis) bringing her fluke down, creating a foot print or fluke print at the ocean surface, with streams of bubbles trailing from the leading points...
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90835953 A female Humpback whale (Megaptera novaeangliae australis) bringing her fluke down, creating a foot print or fluke print at the ocean surface, with streams of bubbles trailing from the leading points...© Tony Wu
Ecoguards / rangers on trail through, Salonga National Park, Democratic Republic of Congo. May 2017.
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90833524 Ecoguards / rangers on trail through, Salonga National Park, Democratic Republic of Congo. May 2017.© Karine Aigner
Aerial view of European flamingos (Phoenicopterus roseus) and their trails as they forage for food, Camargue France. September.
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90831187 Aerial view of European flamingos (Phoenicopterus roseus) and their trails as they forage for food, Camargue France. September.© Jean E. Roche
Ashy mining bee (Andrena cineraria) male feeding on Dandelion (Taraxacum officinale) Monmouthshire, Wales, UK.
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90823243 Ashy mining bee (Andrena cineraria) male feeding on Dandelion (Taraxacum officinale) Monmouthshire, Wales, UK.© Phil Savoie
Bengal Tiger (Panthera tigris) foot print / pugmark on wet sand, Tadoba National Park, India
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90820059 Bengal Tiger (Panthera tigris) foot print / pugmark on wet sand, Tadoba National Park, India© Yashpal Rathore
Ural owl (Strix uralensis) print in snow after landing on prey, Valgamaa county, Southern Estonia. January
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90818120 Ural owl (Strix uralensis) print in snow after landing on prey, Valgamaa county, Southern Estonia. January© Sven Zacek
Leopard (Panthera pardus) female walking in a gully, South Luangwa Narional Park, Zambia. August
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90807276 Leopard (Panthera pardus) female walking in a gully, South Luangwa Narional Park, Zambia. August© Kristel Richard
Wolf (Canis lupus) footprints, Saxony-Anhalt, Germany, August.
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90804057 Wolf (Canis lupus) footprints, Saxony-Anhalt, Germany, August.© Axel Gomille
Wolf (Canis lupus) footprints in tire tracks, Saxony-Anhalt, Germany, August.
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90804056 Wolf (Canis lupus) footprints in tire tracks, Saxony-Anhalt, Germany, August.© Axel Gomille
Wolf (Canis lupus) footprints, Saxony, Germany, July.
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90804054 Wolf (Canis lupus) footprints, Saxony, Germany, July.© Axel Gomille
Wolf (Canis lupus) footprints, Saxony, Germany, August.
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90804051 Wolf (Canis lupus) footprints, Saxony, Germany, August.© Axel Gomille
Humpback whale (Megaptera novaeangliae) trail on water surface from whale bubble-netting. Southeast Alaska, USA. June.
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90801175 Humpback whale (Megaptera novaeangliae) trail on water surface from whale bubble-netting. Southeast Alaska, USA. June.© Doc White
Long legged flies (Dolichopodidae) on puddle in foot print of Long Horn cattle. Knepp Wildland Project, formerly intensive farmland now turned to conservation and sustainable farming. Horsham, West Su...
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90800627 Long legged flies (Dolichopodidae) on puddle in foot print of Long Horn cattle. Knepp Wildland Project, formerly intensive farmland now turned to conservation and sustainable farming. Horsham, West Su...© David Woodfall
Wild Apennine wolf (Canis lupus italicus) tracks in snow. Central Apennines, Abruzzo, Italy. February.
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90799479 Wild Apennine wolf (Canis lupus italicus) tracks in snow. Central Apennines, Abruzzo, Italy. February.© Bruno D'Amicis
Wolf (Canis lupus) paw print, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
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90790339 Wolf (Canis lupus) paw print, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia© Sergey Gorshkov
Derwent Water at Keswick in the Lake District completely frozen over during the December 2010 big chill, with footsteps from a walker who walked across the lake. December 2010
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90789258 Derwent Water at Keswick in the Lake District completely frozen over during the December 2010 big chill, with footsteps from a walker who walked across the lake. December 2010© Ashley Cooper
Footprint in snow, Ambleside, Lake District, Cumbria, England, UK. February 2009
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90789257 Footprint in snow, Ambleside, Lake District, Cumbria, England, UK. February 2009© Ashley Cooper
Polar Bear footprints in melting permafrost on Northern Svalbard. July 2013
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90788546 Polar Bear footprints in melting permafrost on Northern Svalbard. July 2013© Ashley Cooper
Frozen hand print on an iced over car window, Ambleside, Cumbria UK. December 2008
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90788423 Frozen hand print on an iced over car window, Ambleside, Cumbria UK. December 2008© Ashley Cooper
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