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    • Paternal Behaviour
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Tiger cardinalfish (Cheilodipterus macrodon) male mouth brooding a clutch of eggs behind his sharp teeth. Gubal Island, Egypt. Strait Of Gubal, Red Sea.
90802301
90802301 Tiger cardinalfish (Cheilodipterus macrodon) male mouth brooding a clutch of eggs behind his sharp teeth. Gubal Island, Egypt. Strait Of Gubal, Red Sea.© Alex Mustard
Lesser rhea (Pterocnemia pennata) with chicks under wings Valdes Peninsula, Chubut, Patagonia, Argentina  -  Gabriel Rojo/ npl
90672430
90672430 Lesser rhea (Pterocnemia pennata) with chicks under wings Valdes Peninsula, Chubut, Patagonia, Argentina© Gabriel Rojo / NPL
Lesser rhea (Pterocnemia pennata) with chicks under wings Valdes Peninsula, Chubut, Patagonia, Argentina  -  Gabriel Rojo/ npl
90672429
90672429 Lesser rhea (Pterocnemia pennata) with chicks under wings Valdes Peninsula, Chubut, Patagonia, Argentina© Gabriel Rojo / NPL
Lesser rhea (Pterocnemia pennata) with chicks Valdes Peninsula, Chubut, Patagonia, Argentina  -  Gabriel Rojo/ npl
90672428
90672428 Lesser rhea (Pterocnemia pennata) with chicks Valdes Peninsula, Chubut, Patagonia, Argentina© Gabriel Rojo / NPL
Jawfish (Opistognathus iyonis) male mouth brooding eggs, which are hatching with juveniles swimming away. Yamaguchi Prefecture, Honshu, Japan. July.
90854057
90854057 Jawfish (Opistognathus iyonis) male mouth brooding eggs, which are hatching with juveniles swimming away. Yamaguchi Prefecture, Honshu, Japan. July.©
Male Giant water bug (Belostomatidae family) carrying eggs on back, captive
90847718
90847718 Male Giant water bug (Belostomatidae family) carrying eggs on back, captive©
A mature male Fat greenling (Hexagrammos otakii) or Ainame fish tending to eggs in his territory, Hokkaido, Japan. October.
90835972
90835972 A mature male Fat greenling (Hexagrammos otakii) or Ainame fish tending to eggs in his territory, Hokkaido, Japan. October.© Tony Wu
A mature male Fat greenling (Hexagrammos otakii) or Ainame fish tending to eggs in his territory, Hokkaido, Japan. October.
90835971
90835971 A mature male Fat greenling (Hexagrammos otakii) or Ainame fish tending to eggs in his territory, Hokkaido, Japan. October.© Tony Wu
A mature male Fat greenling (Hexagrammos otakii) or Ainame fish tending to eggs in his territory, Hokkaido, Japan. October.
90835970
90835970 A mature male Fat greenling (Hexagrammos otakii) or Ainame fish tending to eggs in his territory, Hokkaido, Japan. October.© Tony Wu
A mature male Fat greenling (Hexagrammos otakii) or Ainame fish tending to eggs in his territory, Hokkaido, Japan. October.
90835969
90835969 A mature male Fat greenling (Hexagrammos otakii) or Ainame fish tending to eggs in his territory, Hokkaido, Japan. October.© Tony Wu
Chestnut-bellied sandgrouse (Pterocles exustus olivascens) male soaking feathers to provide chicks at nest with water. Meru National Park, Kenya.
90821953
90821953 Chestnut-bellied sandgrouse (Pterocles exustus olivascens) male soaking feathers to provide chicks at nest with water. Meru National Park, Kenya.© Melvin Grey
Smooth lumpsucker (Aptocyclus ventricosus) male watching over brood of eggs. Female dies after egg laying while male dies once the eggs have hatched. Hokkaido, Japan. May.
90821115
90821115 Smooth lumpsucker (Aptocyclus ventricosus) male watching over brood of eggs. Female dies after egg laying while male dies once the eggs have hatched. Hokkaido, Japan. May.© Tony Wu
Spotnape cardinalfish (Ostorhinchus notatus) pair, male taking cluster of fertilised eggs into mouth to brood. Shizuoka Prefecture, Honshu, Japan. June.
90821096
90821096 Spotnape cardinalfish (Ostorhinchus notatus) pair, male taking cluster of fertilised eggs into mouth to brood. Shizuoka Prefecture, Honshu, Japan. June.© Tony Wu
Spotnape cardinalfish (Ostorhinchus notatus) pair, male taking cluster of fertilised eggs into mouth to brood. Shizuoka Prefecture, Honshu, Japan. June.
90821095
90821095 Spotnape cardinalfish (Ostorhinchus notatus) pair, male taking cluster of fertilised eggs into mouth to brood. Shizuoka Prefecture, Honshu, Japan. June.© Tony Wu
Burrowing owl (Athene cunicularia) male, insect in beak to hand over to female to feed to chicks. In evening light, Marana, Arizona, USA.
90819525
90819525 Burrowing owl (Athene cunicularia) male, insect in beak to hand over to female to feed to chicks. In evening light, Marana, Arizona, USA.© Jack Dykinga
Burrowing owl (Athene cunicularia) male covering fledgling with wings to protect from danger. Fledgling vocalising and flapping wings in protest. Marana, Arizona, USA. May. Sequence 2/2.
90819499
90819499 Burrowing owl (Athene cunicularia) male covering fledgling with wings to protect from danger. Fledgling vocalising and flapping wings in protest. Marana, Arizona, USA. May. Sequence 2/2.© Jack Dykinga
Burrowing owl (Athene cunicularia) male covering fledgling with wings to protect from danger. Fledgling flapping wings in protest. Marana, Arizona, USA. May. Sequence 1/2.
90819498
90819498 Burrowing owl (Athene cunicularia) male covering fledgling with wings to protect from danger. Fledgling flapping wings in protest. Marana, Arizona, USA. May. Sequence 1/2.© Jack Dykinga
Burrowing owl (Athene cunicularia) male feeding insect to chicks. Marana, Arizona, USA. May.
90819494
90819494 Burrowing owl (Athene cunicularia) male feeding insect to chicks. Marana, Arizona, USA. May.© Jack Dykinga
Burrowing owl (Athene cunicularia) male flying to feed chicks, insect prey in talon. Marana, Arizona, USA. May.
90819493
90819493 Burrowing owl (Athene cunicularia) male flying to feed chicks, insect prey in talon. Marana, Arizona, USA. May.© Jack Dykinga
Burrowing owl (Athene cunicularia) male flying with insect in beak. Will present prey to female to feed to chicks. In evening light, Marana, Arizona, USA. May.
90819492
90819492 Burrowing owl (Athene cunicularia) male flying with insect in beak. Will present prey to female to feed to chicks. In evening light, Marana, Arizona, USA. May.© Jack Dykinga
Comb-crested jacana (Irediparra gallinacea) male and four eggs. The male incubates the eggs. Corroboree Billabong, Mary River, Northern Territory, Australia.
90819476
90819476 Comb-crested jacana (Irediparra gallinacea) male and four eggs. The male incubates the eggs. Corroboree Billabong, Mary River, Northern Territory, Australia.© Steven David Miller
Ringtailed cardinalfish (Apogon aureus) male mouthbrooding eggs. The colour of these eggs shows that they are newly laid. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.
90812271
90812271 Ringtailed cardinalfish (Apogon aureus) male mouthbrooding eggs. The colour of these eggs shows that they are newly laid. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.© Alex Mustard
Portrait of a male tiger cardinalfish (Cheilodipterus macrodon) holding its eggs in his mouth. Gubal Island, Egypt. Gulf of Suez, Red Sea
90812203
90812203 Portrait of a male tiger cardinalfish (Cheilodipterus macrodon) holding its eggs in his mouth. Gubal Island, Egypt. Gulf of Suez, Red Sea© Alex Mustard
Fleischmann's Glassfrog (Hyalinobatrachium fleischmanni) male attending eggs on underside of leaf. Osa Peninsula, Costa Rica.
90811235
90811235 Fleischmann's Glassfrog (Hyalinobatrachium fleischmanni) male attending eggs on underside of leaf. Osa Peninsula, Costa Rica.© Alex Hyde
Tiger cardinalfish (Cheilodipterus macrodon) male mouth broods a clutch of eggs behind his sharp teeth. Gubal Island, Egypt. Strait Of Gubal, Red Sea.
90802239
90802239 Tiger cardinalfish (Cheilodipterus macrodon) male mouth broods a clutch of eggs behind his sharp teeth. Gubal Island, Egypt. Strait Of Gubal, Red Sea.© Alex Mustard
Capybara male with young (Hydrochoerus hydrochaeris) Pantanal, Mato Grosso do Sul, Brazil
90801049
90801049 Capybara male with young (Hydrochoerus hydrochaeris) Pantanal, Mato Grosso do Sul, Brazil© Franco Banfi
Jabiru stork (Jabiru mycteria) male at the nest, feeding its chicks, Pantanal, Brazil.
90798415
90798415 Jabiru stork (Jabiru mycteria) male at the nest, feeding its chicks, Pantanal, Brazil.© Luke Massey
Jabiru stork (Jabiru mycteria) male at the nest, feeding its chicks, Pantanal, Brazil.
90798414
90798414 Jabiru stork (Jabiru mycteria) male at the nest, feeding its chicks, Pantanal, Brazil.© Luke Massey
Jabiru stork (Jabiru mycteria) male at the nest with its chicks, Pantanal, Brazil.
90798406
90798406 Jabiru stork (Jabiru mycteria) male at the nest with its chicks, Pantanal, Brazil.© Luke Massey
Jabiru stork (Jabiru mycteria) male at the nest with its chicks, Pantanal, Brazil.
90798405
90798405 Jabiru stork (Jabiru mycteria) male at the nest with its chicks, Pantanal, Brazil.© Luke Massey
Rhea (Rhea americana) male with his chicks, Pantanal, Brazil.
90798391
90798391 Rhea (Rhea americana) male with his chicks, Pantanal, Brazil.© Luke Massey
Greylag goose (Anser anser) family, two adults and two goslings on water, Danube Delta, Romania. May.
90797579
90797579 Greylag goose (Anser anser) family, two adults and two goslings on water, Danube Delta, Romania. May.© Loic Poidevin
Straw-coloured fruit bat (Eidolon helvum), female flying carrying young on front. Lamin, Gambia.
90795746
90795746 Straw-coloured fruit bat (Eidolon helvum), female flying carrying young on front. Lamin, Gambia.© Bernard Castelein
Ringtailed cardinalfish (Apogon aureus) male mouthbrooding eggs. The colour of these eggs shows that they are close to hatching. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific...
90787561
90787561 Ringtailed cardinalfish (Apogon aureus) male mouthbrooding eggs. The colour of these eggs shows that they are close to hatching. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific...© Alex Mustard
Emu (Dromaius novaehollandiae) male with chicks, Victoria, Australia
90786394
90786394 Emu (Dromaius novaehollandiae) male with chicks, Victoria, Australia© Dave Watts
Emu (Dromaius novaehollandiae) male with chicks, Victoria, Australia
90786389
90786389 Emu (Dromaius novaehollandiae) male with chicks, Victoria, Australia© Dave Watts
Emu (Dromaius novaehollandiae) male with chicks, Victoria, Australia
90786388
90786388 Emu (Dromaius novaehollandiae) male with chicks, Victoria, Australia© Dave Watts
Ostrich (Struthio camelus massaicus) male sitting , guarding eggs. Kenya.
90779126
90779126 Ostrich (Struthio camelus massaicus) male sitting , guarding eggs. Kenya.© Enrique Lopez-Tapia
Ostrich (Struthio camelus massaicus) male sitting, guarding eggs. Kenya.
90779125
90779125 Ostrich (Struthio camelus massaicus) male sitting, guarding eggs. Kenya.© Enrique Lopez-Tapia
False clown anemonefish (Amphiprion oceallaris) male tending  his eggs, cleaning them with his mouth and aerating them by beating his pectoral fins. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, T...
90768075
90768075 False clown anemonefish (Amphiprion oceallaris) male tending his eggs, cleaning them with his mouth and aerating them by beating his pectoral fins. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, T...© Alex Mustard
Hairchin goby (Sagamia geneionema) male measuring about 5cm in length. This fish was guarding eggs that had been laid by a female. Miho, Shizuoka Prefecture, Japan, Pacific.
90762414
90762414 Hairchin goby (Sagamia geneionema) male measuring about 5cm in length. This fish was guarding eggs that had been laid by a female. Miho, Shizuoka Prefecture, Japan, Pacific.© Tony Wu
Hairchin goby (Sagamia geneionema) male protecting a brood of eggs. Miho, Shizuoka Prefecture, Japan, Pacific.
90762413
90762413 Hairchin goby (Sagamia geneionema) male protecting a brood of eggs. Miho, Shizuoka Prefecture, Japan, Pacific.© Tony Wu
Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic. Vulnerable species
90753584
90753584 Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic. Vulnerable species© Yashpal Rathore
Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic.
90753583
90753583 Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic.© Yashpal Rathore
Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic.
90753582
90753582 Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic.© Yashpal Rathore
Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic.
90753581
90753581 Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic.© Yashpal Rathore
Humayuns night frog (Nyctibatrachus humayuni), male guarding cluster of eggs laid on rock near stream. Western Ghats Amboli, Maharashtra, India. Endemic. Vulnerable species
90753574
90753574 Humayuns night frog (Nyctibatrachus humayuni), male guarding cluster of eggs laid on rock near stream. Western Ghats Amboli, Maharashtra, India. Endemic. Vulnerable species© Yashpal Rathore
Humayuns night frog (Nyctibatrachus humayuni), male guarding cluster of eggs laid on rock near stream. Western Ghats Amboli, Maharashtra, India. Endemic. Vulnerable species
90753573
90753573 Humayuns night frog (Nyctibatrachus humayuni), male guarding cluster of eggs laid on rock near stream. Western Ghats Amboli, Maharashtra, India. Endemic. Vulnerable species© Yashpal Rathore
African lion (Panthera leo) male with tiny cub, Masai Mara, Kenya.
90726720
90726720 African lion (Panthera leo) male with tiny cub, Masai Mara, Kenya.© Andy Rouse
Common kingfisher (Alcedo atthis) male flying to nest with fish, Lorraine, France, May
90720839
90720839 Common kingfisher (Alcedo atthis) male flying to nest with fish, Lorraine, France, May© Michel Poinsignon
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