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    • Knuckle Walking
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Chimpanzee (Pan troglodytes) alpha male 'Kris' knucklewalking across log, endangered, Gombe Stream National Park, Tanzania  -  Ingo Arndt
00762316
00762316 Chimpanzee (Pan troglodytes) alpha male 'Kris' knucklewalking across log, endangered, Gombe Stream National Park, Tanzania© Ingo Arndt
Chimpanzee (Pan troglodytes) knuckle-walking through grass, La Vallee Des Singes Primate Center, France  -  Cyril Ruoso
00620532
00620532 Chimpanzee (Pan troglodytes) knuckle-walking through grass, La Vallee Des Singes Primate Center, France© Cyril Ruoso
Bonobo (Pan paniscus) knuckle-walking through grass, La Vallee Des Singes Primate Center, France  -  Cyril Ruoso
00620529
00620529 Bonobo (Pan paniscus) knuckle-walking through grass, La Vallee Des Singes Primate Center, France© Cyril Ruoso
Bonobo (Pan paniscus) knuckle-walking through grass, native to Africa  -  Cyril Ruoso
00620525
00620525 Bonobo (Pan paniscus) knuckle-walking through grass, native to Africa© Cyril Ruoso
Chimpanzee (Pan troglodytes) pair knuckle-walking, La Vallee Des Singes Primate Center, France  -  Cyril Ruoso
00620518
00620518 Chimpanzee (Pan troglodytes) pair knuckle-walking, La Vallee Des Singes Primate Center, France© Cyril Ruoso
Eastern Chimpanzee (Pan troglodytes schweinfurthii) five year old infant male, named Google, Gombe National Park, Tanzania
00598460
00598460 Eastern Chimpanzee (Pan troglodytes schweinfurthii) five year old infant male, named Google, Gombe National Park, Tanzania© Anup Shah
Eastern Chimpanzee (Pan troglodytes schweinfurthii) male, eighteen years old, followed by troop, Gombe National Park, Tanzania
00567418
00567418 Eastern Chimpanzee (Pan troglodytes schweinfurthii) male, eighteen years old, followed by troop, Gombe National Park, Tanzania© Anup Shah
Chimpanzee (Pan troglodytes) mother carrying young, Singapore Zoo, Singapore
00565507
00565507 Chimpanzee (Pan troglodytes) mother carrying young, Singapore Zoo, Singapore© Juergen & Christine Sohns
Chimpanzee (Pan troglodytes) sub-adult running, Singapore Zoo, Singapore
00565499
00565499 Chimpanzee (Pan troglodytes) sub-adult running, Singapore Zoo, Singapore© Juergen & Christine Sohns
Eastern Chimpanzee (Pan troglodytes schweinfurthii) males walking along forest floor, Gombe National Park, Tanzania
00553757
00553757 Eastern Chimpanzee (Pan troglodytes schweinfurthii) males walking along forest floor, Gombe National Park, Tanzania© Anup Shah
Eastern Chimpanzee (Pan troglodytes schweinfurthii) infant male, one year old, running, Gombe National Park, Tanzania
00553719
00553719 Eastern Chimpanzee (Pan troglodytes schweinfurthii) infant male, one year old, running, Gombe National Park, Tanzania© Fiona Rogers
Sumatran Orangutan (Pongo abelii) sub-adult male, Gunung Leuser National Park, north Sumatra, Indonesia  -  Suzi Eszterhas
00443967
00443967 Sumatran Orangutan (Pongo abelii) sub-adult male, Gunung Leuser National Park, north Sumatra, Indonesia© Suzi Eszterhas
Chimpanzee (Pan troglodytes) male running, western Uganda  -  Suzi Eszterhas
00438355
00438355 Chimpanzee (Pan troglodytes) male running, western Uganda© Suzi Eszterhas
Chimpanzee (Pan troglodytes) group, rescued individuals moving through entrance to sleeping area, Ngamba Island Chimpanzee Sanctuary, Uganda  -  Suzi Eszterhas
00438026
00438026 Chimpanzee (Pan troglodytes) group, rescued individuals moving through entrance to sleeping area, Ngamba Island Chimpanzee Sanctuary, Uganda© Suzi Eszterhas
Mountain Gorilla (Gorilla gorilla beringei) male, Virunga Mountains, Rwanda  -  Gerry Ellis
00205163
00205163 Mountain Gorilla (Gorilla gorilla beringei) male, Virunga Mountains, Rwanda© Gerry Ellis
Eastern Chimpanzee (Pan troglodytes schweinfurthii) walking on knuckles, Mahale Mountains, Tanzania  -  Ferrero-Labat/ Auscape
00441893
00441893 Eastern Chimpanzee (Pan troglodytes schweinfurthii) walking on knuckles, Mahale Mountains, Tanzania© Ferrero-Labat/ Auscape
Mountain Gorilla (Gorilla beringei beringei) 'silverback' alpha male, leading family group into clearing, Volcanoes National Park, Virunga Mountains, Rwanda, June  -  Martin Hale/ FLPA
80163236
80163236 Mountain Gorilla (Gorilla beringei beringei) 'silverback' alpha male, leading family group into clearing, Volcanoes National Park, Virunga Mountains, Rwanda, June© Martin Hale / FLPA
Mountain Gorilla (Gorilla beringei beringei) 'silverback' alpha male, leading family group into clearing, Volcanoes National Park, Virunga Mountains, Rwanda, June  -  Martin Hale/ FLPA
80163233
80163233 Mountain Gorilla (Gorilla beringei beringei) 'silverback' alpha male, leading family group into clearing, Volcanoes National Park, Virunga Mountains, Rwanda, June© Martin Hale / FLPA
Eastern Chimpanzee (Pan troglodytes schweinfurthii) twenty-five year old male, named Faustino, leading troop, Gombe National Park, Tanzania
00598474
00598474 Eastern Chimpanzee (Pan troglodytes schweinfurthii) twenty-five year old male, named Faustino, leading troop, Gombe National Park, Tanzania© Anup Shah
Eastern Chimpanzee (Pan troglodytes schweinfurthii) twelve year old male, named Tom, running, Gombe National Park, Tanzania
00581511
00581511 Eastern Chimpanzee (Pan troglodytes schweinfurthii) twelve year old male, named Tom, running, Gombe National Park, Tanzania© Anup Shah
Eastern Chimpanzee (Pan troglodytes schweinfurthii) seventeen year old male, named Sampson, calling, Gombe National Park, Tanzania
00581510
00581510 Eastern Chimpanzee (Pan troglodytes schweinfurthii) seventeen year old male, named Sampson, calling, Gombe National Park, Tanzania© Anup Shah
Eastern Chimpanzee (Pan troglodytes schweinfurthii) nineteen year old male, named Titan, Gombe National Park, Tanzania
00581484
00581484 Eastern Chimpanzee (Pan troglodytes schweinfurthii) nineteen year old male, named Titan, Gombe National Park, Tanzania© Anup Shah
Eastern Chimpanzee (Pan troglodytes schweinfurthii) nine year old male, Named Gimli, Gombe National Park, Tanzania
00581481
00581481 Eastern Chimpanzee (Pan troglodytes schweinfurthii) nine year old male, Named Gimli, Gombe National Park, Tanzania© Anup Shah
Chimpanzee (Pan troglodytes) orphan Larry knucklwalking, Ape Action Africa, Mefou Primate Sanctuary, Cameroon
00573747
00573747 Chimpanzee (Pan troglodytes) orphan Larry knucklwalking, Ape Action Africa, Mefou Primate Sanctuary, Cameroon© Gerry Ellis
Eastern Chimpanzee (Pan troglodytes schweinfurthii) male, thirty-four years old, Gombe National Park, Tanzania
00567430
00567430 Eastern Chimpanzee (Pan troglodytes schweinfurthii) male, thirty-four years old, Gombe National Park, Tanzania© Anup Shah
Eastern chimpanzee (Pan troglodytes schweinfurtheii) males knuckle walking along the shore of Lake Tanganyika . Gombe National Park, Tanzania. May 2014.
90825155
90825155 Eastern chimpanzee (Pan troglodytes schweinfurtheii) males knuckle walking along the shore of Lake Tanganyika . Gombe National Park, Tanzania. May 2014.© Anup Shah
Eastern Chimpanzee (Pan troglodytes schweinfurthii) knuckle walking up hillside, Gombe National Park, Tanzania  -  Anup Shah/ npl
00414927
00414927 Eastern Chimpanzee (Pan troglodytes schweinfurthii) knuckle walking up hillside, Gombe National Park, Tanzania© Anup Shah/ npl
Western Lowland Gorilla (Gorilla gorilla gorilla) silverback male knucklewalking, endangered, Odzala National Park, Democratic Republic of the Congo  -  Jabruson/ npl
00413668
00413668 Western Lowland Gorilla (Gorilla gorilla gorilla) silverback male knucklewalking, endangered, Odzala National Park, Democratic Republic of the Congo© Jabruson / NPL
Orangutan (Pongo pygmaeus) four year old knucklewalking, Tanjung Puting National Park, Borneo, Indonesia  -  Suzi Eszterhas
00553605
00553605 Orangutan (Pongo pygmaeus) four year old knucklewalking, Tanjung Puting National Park, Borneo, Indonesia© Suzi Eszterhas
Orangutan (Pongo pygmaeus) mother and two year old young, Tanjung Puting National Park, Borneo, Indonesia  -  Suzi Eszterhas
00553598
00553598 Orangutan (Pongo pygmaeus) mother and two year old young, Tanjung Puting National Park, Borneo, Indonesia© Suzi Eszterhas
Bonobo (Pan paniscus) female 'Nioki' carrying her baby 'Bomango' aged 10 months on her back, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo October  -  Anup Shah/ npl
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90355470 Bonobo (Pan paniscus) female 'Nioki' carrying her baby 'Bomango' aged 10 months on her back, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo October© Anup Shah / NPL
Bonobo (Pan paniscus) male 'Kikongo' walking through the forest, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo October  -  Anup Shah/ npl
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90355443 Bonobo (Pan paniscus) male 'Kikongo' walking through the forest, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo October© Anup Shah / NPL
Bonobo (Pan paniscus) female 'Opala' carrying her juvenile son 'Pole' aged 4 years on her back, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo October  -  Anup Shah/ npl
90355392
90355392 Bonobo (Pan paniscus) female 'Opala' carrying her juvenile son 'Pole' aged 4 years on her back, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo October© Anup Shah / NPL
Bonobo (Pan paniscus) female walking, carrying her baby under her belly, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo October  -  Anup Shah/ npl
90355253
90355253 Bonobo (Pan paniscus) female walking, carrying her baby under her belly, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo October© Anup Shah / NPL
Bonobo (Pan paniscus) mature male 'Kasongo' walking at the edge of a lake, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo October  -  Fiona Rogers/ npl
90354884
90354884 Bonobo (Pan paniscus) mature male 'Kasongo' walking at the edge of a lake, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo October© Fiona Rogers / NPL
Bonobo (Pan paniscus) adolescent male, standing portrait, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo October  -  Fiona Rogers/ npl
90354870
90354870 Bonobo (Pan paniscus) adolescent male, standing portrait, Lola Ya Bonobo Sanctuary, Democratic Republic of Congo October© Fiona Rogers / NPL
Subadult male Chimpanzee (Pan troglodytes schweinfurthii) Hawa (13 years) Budongo Forest Reserve, Masindi, Uganda, Africa December  -  Florian Mollers/ npl
90225839
90225839 Subadult male Chimpanzee (Pan troglodytes schweinfurthii) Hawa (13 years) Budongo Forest Reserve, Masindi, Uganda, Africa December© Florian Mllers / NPL
Subadult male Chimpanzee (Pan troglodytes schweinfurthii) Hawa (13 years) Budongo Forest Reserve, Masindi, Uganda, Africa December  -  Florian Mollers/ npl
90225833
90225833 Subadult male Chimpanzee (Pan troglodytes schweinfurthii) Hawa (13 years) Budongo Forest Reserve, Masindi, Uganda, Africa December© Florian Mllers / NPL
Male Chimpanzees (Pan troglodytes schweinfurthii) Maani (front) and Kato approaching in impressive posture, zoom-effect Budongo Forest Reserve, Masindi, Uganda, Africa December  -  Florian Mollers/ npl
90225828
90225828 Male Chimpanzees (Pan troglodytes schweinfurthii) Maani (front) and Kato approaching in impressive posture, zoom-effect Budongo Forest Reserve, Masindi, Uganda, Africa December© Florian Mllers / NPL
Chimpanzee (Pan troglodytes schweinfurthii) mother Nambi crossing forest clearing carrying juvenile Night (3 years) on her back Budongo Forest Reserve, Masindi, Uganda, Africa December  -  Florian Mollers/ npl
90225677
90225677 Chimpanzee (Pan troglodytes schweinfurthii) mother Nambi crossing forest clearing carrying juvenile Night (3 years) on her back Budongo Forest Reserve, Masindi, Uganda, Africa December© Florian Mllers / NPL
Subadult Chimpanzee (Pan troglodytes schweinfurthii) male Musa crossing rainforest clearing carrying figs in his mouth Budongo Forest Reserve, Masindi, Uganda, Africa December  -  Florian Mollers/ npl
90225593
90225593 Subadult Chimpanzee (Pan troglodytes schweinfurthii) male Musa crossing rainforest clearing carrying figs in his mouth Budongo Forest Reserve, Masindi, Uganda, Africa December© Florian Mllers / NPL
Male Chimpanzee running, Kenya  -  Karl Ammann/ npl
90209996
90209996 Male Chimpanzee running, Kenya© Karl Ammann / NPL
Western lowland gorilla (Gorilla gorilla gorilla) silverback knuckle walking, Lokoue Bai, Odzala NP, Congo  -  Jabruson/ npl
90209128
90209128 Western lowland gorilla (Gorilla gorilla gorilla) silverback knuckle walking, Lokoue Bai, Odzala NP, Congo© Jabruson / NPL
Western lowland gorilla (Gorilla gorilla gorilla) Silverback male knuckle walking, Lokoue Bai, Odzala NP, Congo Rep  -  Jabruson/ npl
90201401
90201401 Western lowland gorilla (Gorilla gorilla gorilla) Silverback male knuckle walking, Lokoue Bai, Odzala NP, Congo Rep© Jabruson / NPL
Chimpanzee (Pan troglodytes) juvenile knuckle walking, captive, Chimfunshi wildlife orphanage, Zambia  -  David Pike/ npl
90162742
90162742 Chimpanzee (Pan troglodytes) juvenile knuckle walking, captive, Chimfunshi wildlife orphanage, Zambia© David Pike / NPL
Western lowland gorilla (Gorilla gorilla gorilla) juvenile knuckle walking, Lokoue Bai, Odzala NP, Congo Rep  -  Jabruson/ npl
90161615
90161615 Western lowland gorilla (Gorilla gorilla gorilla) juvenile knuckle walking, Lokoue Bai, Odzala NP, Congo Rep© Jabruson / NPL
Western lowland gorilla (Gorilla gorilla gorilla) Juvenile male knuckle walking, Lokoue Bai, Odzala NP, Congo Rep  -  Jabruson/ npl
90161614
90161614 Western lowland gorilla (Gorilla gorilla gorilla) Juvenile male knuckle walking, Lokoue Bai, Odzala NP, Congo Rep© Jabruson / NPL
Male Western lowland gorilla walking, Lokoue Bai (Gorilla g gorilla) Odzala NP, Rep Congo  -  Jabruson/ npl
90160126
90160126 Male Western lowland gorilla walking, Lokoue Bai (Gorilla g gorilla) Odzala NP, Rep Congo© Jabruson / NPL
Male silverback Mountain gorilla (Gorilla gorilla beringei) knuckle walking, Volcanoes National Park, Rwanda, Africa  -  Eric Baccega/ npl
90145381
90145381 Male silverback Mountain gorilla (Gorilla gorilla beringei) knuckle walking, Volcanoes National Park, Rwanda, Africa© Eric Baccega / NPL
Female Chimpanzee 'Patti' (39-years-old) carries jockey-riding offspring 'Tarzan' on her back (Pan troglodytes schweinfurtheii), Kasekela community, Gombe NP, Tanzania 2002  -  Anup Shah/ npl
90142037
90142037 Female Chimpanzee 'Patti' (39-years-old) carries jockey-riding offspring 'Tarzan' on her back (Pan troglodytes schweinfurtheii), Kasekela community, Gombe NP, Tanzania 2002© Anup Shah / NPL
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