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Gentoo penguins (Pygoscelis papua) calling and defending their eggs from Southern Skua (Stercorarius antarcticus) hunting for unattended eggs or chicks, Sea Lion island, The Falklands, South Atlantic.
90699531
90699531 Gentoo penguins (Pygoscelis papua) calling and defending their eggs from Southern Skua (Stercorarius antarcticus) hunting for unattended eggs or chicks, Sea Lion island, The Falklands, South Atlantic.© Michael Pitts
Great frigate birds (Fregata minor) sitting in tree, Aldabra Atoll, Seychelles, Indian Ocean
90698568
90698568 Great frigate birds (Fregata minor) sitting in tree, Aldabra Atoll, Seychelles, Indian Ocean© Cheryl-Samantha Owen / naturepl
Two male urban Red foxes, mouths open,  one in submissive posture {Vulpes vulpes} London, UK  -  Laurent Geslin/ npl
90212835
90212835 Two male urban Red foxes, mouths open, one in submissive posture {Vulpes vulpes} London, UK© Laurent Geslin / NPL
Two dominant male urban Red foxes fighting on road, mouths open {Vulpes vulpes} London, UK  -  Laurent Geslin/ npl
90197680
90197680 Two dominant male urban Red foxes fighting on road, mouths open {Vulpes vulpes} London, UK© Laurent Geslin / NPL
Red deer stag calling in rut {Cervus elaphus} with hinds nearby, Richmond Park, London, UK  -  David Kjaer/ npl
90160135
90160135 Red deer stag calling in rut {Cervus elaphus} with hinds nearby, Richmond Park, London, UK© David Kjaer / NPL
Two Northern elephant seal (Mirounga angustirostris) males fighting for dominance, California, USA  -  Ingo Arndt/ npl
90142095
90142095 Two Northern elephant seal (Mirounga angustirostris) males fighting for dominance, California, USA© Ingo Arndt / NPL
Chimpanzees gather round alpha male 'Frodo' in tree feeding on dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002  -  Anup Shah/ npl
90141012
90141012 Chimpanzees gather round alpha male 'Frodo' in tree feeding on dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002© Anup Shah / NPL
Purple sandpiper (Calidris maritima) aerial of group in snow,  panoramic, Batsfjord, Norway, February. Digitally stitched image.
90700113
90700113 Purple sandpiper (Calidris maritima) aerial of group in snow, panoramic, Batsfjord, Norway, February. Digitally stitched image.© Pal Hermansen
Flock of Purple sandpipers (Calidris maritima) in snow, aerial view, Batsfjord, Finnmark, Norway, February.
90700112
90700112 Flock of Purple sandpipers (Calidris maritima) in snow, aerial view, Batsfjord, Finnmark, Norway, February.© Pal Hermansen
Wreathed hornbills (Rhyiceros undulatus) roosting in tree, Pakke, Arunachal pradesh, India
90699107
90699107 Wreathed hornbills (Rhyiceros undulatus) roosting in tree, Pakke, Arunachal pradesh, India© Sandesh Kadur
Northern gannet (Morus bassanus) flying above breeding colony, Great Saltee Island, Wexford, Ireland, June
90698577
90698577 Northern gannet (Morus bassanus) flying above breeding colony, Great Saltee Island, Wexford, Ireland, June© Roger Powell / naturepl
Northern gannets (Morus bassanus) flying above their breeding colony, Great Saltee Island, Wexford, Ireland, June
90698576
90698576 Northern gannets (Morus bassanus) flying above their breeding colony, Great Saltee Island, Wexford, Ireland, June© Roger Powell / naturepl
Lined sweetlips fish swims with shoal of Large-eyed jacks. Digitally manipulated image  -  Georgette Douwma/ npl
90210361
90210361 Lined sweetlips fish swims with shoal of Large-eyed jacks. Digitally manipulated image© GEORGETTE DOUWMA / NPL
Male Black grouse {Tetrao tetrix} displaying on lek site. Scotland, UK.  -  Niall Benvie/ npl
90195470
90195470 Male Black grouse {Tetrao tetrix} displaying on lek site. Scotland, UK.© Niall Benvie / NPL
Tufted puffin pair (left) interrupted by individual (rt) {Lunda cirrhata} East Russia  -  Ben Osborne/ npl
90182428
90182428 Tufted puffin pair (left) interrupted by individual (rt) {Lunda cirrhata} East Russia© Ben Osborne / NPL
Silhouette of Carrion Crows {Corvus corone} perching on a dead tree at dawn, Kent, UK  -  David Tipling/ npl
90173610
90173610 Silhouette of Carrion Crows {Corvus corone} perching on a dead tree at dawn, Kent, UK© DAVID TIPLING / NPL
Shaggy pholiota fungus on tree {Pholiota squarrosa} Peak District NP, Derbyshire, UK  -  Geoff Simpson/ npl
90167690
90167690 Shaggy pholiota fungus on tree {Pholiota squarrosa} Peak District NP, Derbyshire, UK© Geoff Simpson / NPL
Male Hippopotamus fighting in Mara river(Hippopotamus amphibius), Masai Mara GR, Kenya  -  Anup Shah/ npl
90140778
90140778 Male Hippopotamus fighting in Mara river(Hippopotamus amphibius), Masai Mara GR, Kenya© Anup Shah / NPL
Domestic Cat {Felis catus} Ginger tom 'Fergus' rebuffs friendly advance of his 11-week son 'Septimus' at their first meeting.  -  Jane Burton/ npl
90030709
90030709 Domestic Cat {Felis catus} Ginger tom 'Fergus' rebuffs friendly advance of his 11-week son 'Septimus' at their first meeting.© Jane Burton/ npl
Ruff (Philomachus pugnax) males fighting for hierarchy, Lapland, Sweden  -  Dominique Halleux/ Biosphoto
00489344
00489344 Ruff (Philomachus pugnax) males fighting for hierarchy, Lapland, Sweden© Dominique Halleux/ Biosphoto
Rocky Mountain Bighorn Sheep (Ovis canadensis) males, horning each other in dominance display Whiskey Mountain Sheep ranger, Wind River Mts near Dubois, Wyoming  -  Jeff Vanuga/ npl
90697885
90697885 Rocky Mountain Bighorn Sheep (Ovis canadensis) males, horning each other in dominance display Whiskey Mountain Sheep ranger, Wind River Mts near Dubois, Wyoming© Jeff Vanuga / NPL
African elephants (Loxodonta africana) two bulls sparring at the waterhole in front of the lodge, Ol Donyo Wuas, Mbirikani Group Ranch, Amboseli-Tsavo ecosystem, Chyulu Hills, Kenya  -  Ann & Steve Toon/ npl
90696247
90696247 African elephants (Loxodonta africana) two bulls sparring at the waterhole in front of the lodge, Ol Donyo Wuas, Mbirikani Group Ranch, Amboseli-Tsavo ecosystem, Chyulu Hills, Kenya© Ann & Steve Toon / NPL
African Elephant (Loxodonta africana), young males playing and pretending to copulate Masai-Mara Game Reserve, Kenya  -  Denis Huot/ npl
90694808
90694808 African Elephant (Loxodonta africana), young males playing and pretending to copulate Masai-Mara Game Reserve, Kenya© Denis-Huot / NPL
Topi (Damaliscus lunatus jimela) two males sparring, Masai Mara National Reserve, Kenya March  -  Anup Shah/ npl
90693640
90693640 Topi (Damaliscus lunatus jimela) two males sparring, Masai Mara National Reserve, Kenya March© Anup Shah / NPL
African Wild Dog (Lycaon pictus) mobbing interaction within a pack, Mana Pools National Park, Zimbabwe October 2012  -  Tony Heald/ npl
90692876
90692876 African Wild Dog (Lycaon pictus) mobbing interaction within a pack, Mana Pools National Park, Zimbabwe October 2012© Tony Heald / NPL
African Wild Dog (Lycaon pictus) submissive behaviour within a pack, Mana Pools National Park, Zimbabwe, October 2012  -  Tony Heald/ npl
90692850
90692850 African Wild Dog (Lycaon pictus) submissive behaviour within a pack, Mana Pools National Park, Zimbabwe, October 2012© Tony Heald / NPL
Bison (Bison bison) two males head to head, Yellowstone NP, Wyoming, USA  -  George Sanker/ npl
90690405
90690405 Bison (Bison bison) two males head to head, Yellowstone NP, Wyoming, USA© George Sanker / NPL
Moose (Alces Alces) males fighting, clashing antlers at rut, Grand Teton NP, Wyoming, USA, October  -  George Sanker/ npl
90690391
90690391 Moose (Alces Alces) males fighting, clashing antlers at rut, Grand Teton NP, Wyoming, USA, October© George Sanker / NPL
Northern mockingbird (Mimus polyglottos) adult fighting with Golden-fronted Woodpecker (Melanerpes aurifrons) female over Texas Prickly Pear Cactus (Opuntia lindheimeri) Dinero, Lake Corpus Christi, S...  -  Rolf Nussbaumer/ npl
90690098
90690098 Northern mockingbird (Mimus polyglottos) adult fighting with Golden-fronted Woodpecker (Melanerpes aurifrons) female over Texas Prickly Pear Cactus (Opuntia lindheimeri) Dinero, Lake Corpus Christi, S...© Rolf Nussbaumer / NPL
Nyala (Tragelaphus angasii) bull dominance display, Mkhuze Game Reserve, South Africa, May  -  Ann & Steve Toon/ npl
90686835
90686835 Nyala (Tragelaphus angasii) bull dominance display, Mkhuze Game Reserve, South Africa, May© Ann & Steve Toon / NPL
Western chimpanzee (Pan troglodytes verus) young males 'Jeje' aged 13 years and 'Peley' aged 12 years displaying to each other, Bossou Forest, Mont Nimba, Guinea January 2011 Sequence 3 of 3  -  Anup Shah/ npl
90684184
90684184 Western chimpanzee (Pan troglodytes verus) young males 'Jeje' aged 13 years and 'Peley' aged 12 years displaying to each other, Bossou Forest, Mont Nimba, Guinea January 2011 Sequence 3 of 3© Anup Shah / NPL
Western chimpanzee (Pan troglodytes verus) young males 'Jeje' aged 13 years and 'Peley' aged 12 years displaying to each other, Bossou Forest, Mont Nimba, Guinea January 2011 Sequence 2 of 3  -  Anup Shah/ npl
90684183
90684183 Western chimpanzee (Pan troglodytes verus) young males 'Jeje' aged 13 years and 'Peley' aged 12 years displaying to each other, Bossou Forest, Mont Nimba, Guinea January 2011 Sequence 2 of 3© Anup Shah / NPL
Black grouse (Tetrao tetrix) two males displaying at lek, Bergslagen, Sweden, April  -  Orsolya Haarberg/ npl
90683367
90683367 Black grouse (Tetrao tetrix) two males displaying at lek, Bergslagen, Sweden, April© Orsolya Haarberg / NPL
Alpine ibex (Capra ibex) large ram standing on ridge in snow, Alps, Gran Paradiso National Park, Italy, October  -  Orsolya Haarberg/ npl
90683325
90683325 Alpine ibex (Capra ibex) large ram standing on ridge in snow, Alps, Gran Paradiso National Park, Italy, October© Orsolya Haarberg / NPL
Alpine ibex (Capra ibex) large ram in deep snow, Alps, Gran Paradiso National Park, Italy, October  -  Orsolya Haarberg/ npl
90683321
90683321 Alpine ibex (Capra ibex) large ram in deep snow, Alps, Gran Paradiso National Park, Italy, October© Orsolya Haarberg / NPL
Black Grouse (Tetrao tetrix) two males fighting, Liminka, Finland, March  -  Markus Varesvuo/ npl
90679872
90679872 Black Grouse (Tetrao tetrix) two males fighting, Liminka, Finland, March© Markus Varesvuo / NPL
Black Grouse (Tetrao tetrix) two males fighting, Liminka, Finland, March  -  Markus Varesvuo/ npl
90679871
90679871 Black Grouse (Tetrao tetrix) two males fighting, Liminka, Finland, March© Markus Varesvuo / NPL
Black Grouse (Tetrao tetrix) two males fighting, Liminka, Finland, March  -  Markus Varesvuo/ npl
90679870
90679870 Black Grouse (Tetrao tetrix) two males fighting, Liminka, Finland, March© Markus Varesvuo / NPL
Semi wild Przewalski horses (Equus ferus przewalskii), stallion biting another stallion to chase it from his herd, Parc du Villaret, Causse Mejean, Lozere, France  -  Eric Baccega/ npl
90679574
90679574 Semi wild Przewalski horses (Equus ferus przewalskii), stallion biting another stallion to chase it from his herd, Parc du Villaret, Causse Mejean, Lozere, France© Eric Baccega / NPL
Siamese fighting fish (Betta splendens) two males in lateral display, captive, freshwater, from Malaysia and Thailand  -  Jane Burton/ npl
90679512
90679512 Siamese fighting fish (Betta splendens) two males in lateral display, captive, freshwater, from Malaysia and Thailand© Jane Burton / NPL
Male Red hartebeest (Alcelaphus buselaphus) fighting, Kgalagadi Transfrontier Park, Kalahari desert, South Africa  -  Jouan & Rius/ npl
90679152
90679152 Male Red hartebeest (Alcelaphus buselaphus) fighting, Kgalagadi Transfrontier Park, Kalahari desert, South Africa© Jouan & Rius / NPL
Wild horses, mustangs, in Pryor Mountains, Montana, USA, two stallions face off, one rearing  -  Carol Walker/ npl
90679118
90679118 Wild horses, mustangs, in Pryor Mountains, Montana, USA, two stallions face off, one rearing© Carol Walker / NPL
Northern elephant seal males fighting, California, USA (Mirounga angustirostris) California, USA  -  Ingo Arndt/ npl
90677068
90677068 Northern elephant seal males fighting, California, USA (Mirounga angustirostris) California, USA© Ingo Arndt / NPL
Male Prairie chicken (Tympanuchus cupido) displaying Buena Vista Marsh, Wisconsin, USA  -  Thomas Lazar/ npl
90676114
90676114 Male Prairie chicken (Tympanuchus cupido) displaying Buena Vista Marsh, Wisconsin, USA© Thomas Lazar / NPL
Long-tailed macaque (Macaca fascicularis) chattering in submission to be allowed to groom Bako National Park, Sarawak, Borneo, Malaysia  -  Anup Shah/ npl
90404627
90404627 Long-tailed macaque (Macaca fascicularis) chattering in submission to be allowed to groom Bako National Park, Sarawak, Borneo, Malaysia© Anup Shah / NPL
Long-tailed macaque (Macaca fascicularis) female demonstrating begging gesture to be allowed to groom another Bako National Park, Sarawak, Borneo, Malaysia Mar 2010  -  Anup Shah/ npl
90404562
90404562 Long-tailed macaque (Macaca fascicularis) female demonstrating begging gesture to be allowed to groom another Bako National Park, Sarawak, Borneo, Malaysia Mar 2010© Anup Shah / NPL
Rocky Mountain Bighorn Sheep (Ovis canadensis) young ram in subservient behaviour to dominant ram, Rocky Mountain National Park, Colorado, USA, August  -  Charlie Summers/ npl
90397029
90397029 Rocky Mountain Bighorn Sheep (Ovis canadensis) young ram in subservient behaviour to dominant ram, Rocky Mountain National Park, Colorado, USA, August© Charlie Summers / NPL
Chimpanzee (Pan troglodytes) large alpha male in tropical forest, Western Uganda  -  Suzi Eszterhas/ npl
90387229
90387229 Chimpanzee (Pan troglodytes) large alpha male in tropical forest, Western Uganda© Suzi Eszterhas / NPL
Fallow Deer (Dama dama) bucks fighting at Attingham Park, National Trust, Shropshire, UK, November  -  Will Watson/ npl
90387157
90387157 Fallow Deer (Dama dama) bucks fighting at Attingham Park, National Trust, Shropshire, UK, November© Will Watson / NPL
Coot (Fulica atra) one coot is chasing another one in a territorial dispute, La Dombes lake area, France, April  -  David Pattyn/ npl
90386670
90386670 Coot (Fulica atra) one coot is chasing another one in a territorial dispute, La Dombes lake area, France, April© David Pattyn / NPL
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