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    • Ecosystem Engineer
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Tree felled by beavers,with marks of their teeth on it, near a beaver lodge, underwater. Vieille Thielle (Alte Zihl), Neuchatel, Switzerland. April. Photographed for The Freshwater Project.
90836459
90836459 Tree felled by beavers,with marks of their teeth on it, near a beaver lodge, underwater. Vieille Thielle (Alte Zihl), Neuchatel, Switzerland. April. Photographed for The Freshwater Project.© Michel Roggo
Alcedo giant tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island. This is where the largest extant population exists, acting as ecosystem engineers by maintaining open meadows and digg...
90836176
90836176 Alcedo giant tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island. This is where the largest extant population exists, acting as ecosystem engineers by maintaining open meadows and digg...© Tui De Roy
Alcedo giant tortoises (Chelonoidis vandenburghi) wallowing, Alcedo Volcano, Isabela Island. This is where the largest population still exists, acting as ecosystem engineers by maintaining open meadow...
90836173
90836173 Alcedo giant tortoises (Chelonoidis vandenburghi) wallowing, Alcedo Volcano, Isabela Island. This is where the largest population still exists, acting as ecosystem engineers by maintaining open meadow...© Tui De Roy
Trees felled by beavers, with marks of the teeth, near a Beaver lodge, under water, Vieille Thielle (Alte Zihl), close to Cressier, Canton of Neuchatel, Switzerland. April. Photographed for The Freshw...
90827134
90827134 Trees felled by beavers, with marks of the teeth, near a Beaver lodge, under water, Vieille Thielle (Alte Zihl), close to Cressier, Canton of Neuchatel, Switzerland. April. Photographed for The Freshw...© Michel Roggo
North American  Beaver (Castor canadensis) dam, Northern Rockies, Muskwa-Kechika Protected Area, British Columbia, Canada, July 2011. Photographed for The Freshwater Project
90776192
90776192 North American Beaver (Castor canadensis) dam, Northern Rockies, Muskwa-Kechika Protected Area, British Columbia, Canada, July 2011. Photographed for The Freshwater Project© Michel Roggo
Sockeye Salmon (Oncorhynchus nerka) on spawning ground in groundwater channel, with trees cut by beavers Adams river, British Columbia, Canada, October.    Photographed for the Freshwater Project.
90776186
90776186 Sockeye Salmon (Oncorhynchus nerka) on spawning ground in groundwater channel, with trees cut by beavers Adams river, British Columbia, Canada, October. Photographed for the Freshwater Project.© Michel Roggo
Eurasian beaver (Castor fiber albicus) tree damage, Spessart, Germany, November.
90764358
90764358 Eurasian beaver (Castor fiber albicus) tree damage, Spessart, Germany, November.© Ingo Arndt
Silhouetted Red deer (Cervus elaphus) male calling at dusk, Scotland, UK.
90752987
90752987 Silhouetted Red deer (Cervus elaphus) male calling at dusk, Scotland, UK.© SCOTLAND: The Big Picture
Alder (Alnus glutinosa) tree felled by Eurasian beaver (Castor fiber), Knapdale Forest, Argyll, Scotland, UK, June 2016.
90752977
90752977 Alder (Alnus glutinosa) tree felled by Eurasian beaver (Castor fiber), Knapdale Forest, Argyll, Scotland, UK, June 2016.© SCOTLAND: The Big Picture
Alder (Alnus glutinosa) tree felled by Eurasian beaver (Castor fiber), Knapdale Forest, Argyll, Scotland, UK, June 2016.
90752976
90752976 Alder (Alnus glutinosa) tree felled by Eurasian beaver (Castor fiber), Knapdale Forest, Argyll, Scotland, UK, June 2016.© SCOTLAND: The Big Picture
Alder (Alnus glutinosa) tree felled by Eurasian beaver (Castor fiber), Knapdale Forest, Argyll, Scotland, UK, June 2016.
90752975
90752975 Alder (Alnus glutinosa) tree felled by Eurasian beaver (Castor fiber), Knapdale Forest, Argyll, Scotland, UK, June 2016.© SCOTLAND: The Big Picture
Tree with signs of Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, Argyll, Scotland, UK, June.
90752774
90752774 Tree with signs of Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, Argyll, Scotland, UK, June.© SCOTLAND: The Big Picture
Eurasian beaver (Castor fiber) lodge, Knapdale Forest, Argyll, Scotland, UK, June.
90752770
90752770 Eurasian beaver (Castor fiber) lodge, Knapdale Forest, Argyll, Scotland, UK, June.© SCOTLAND: The Big Picture
Felled tree, a sign of Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, Argyll, Scotland, UK, June.
90752769
90752769 Felled tree, a sign of Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, Argyll, Scotland, UK, June.© SCOTLAND: The Big Picture
Felled tree, a sign of Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, Argyll, Scotland, UK, June.
90752768
90752768 Felled tree, a sign of Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, Argyll, Scotland, UK, June.© SCOTLAND: The Big Picture
Woodland habitat surrounding Eurasian beaver (Castor fiber) lochs showing diversity of vegetation and structure, Knapdale Forest, site of Scottish Beaver Trial, Argyll, Scotland, UK, June.
90752767
90752767 Woodland habitat surrounding Eurasian beaver (Castor fiber) lochs showing diversity of vegetation and structure, Knapdale Forest, site of Scottish Beaver Trial, Argyll, Scotland, UK, June.© SCOTLAND: The Big Picture
Felled tree with signs of Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, Argyll, Scotland, UK, June.
90752766
90752766 Felled tree with signs of Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, Argyll, Scotland, UK, June.© SCOTLAND: The Big Picture
Eurasian beaver (Castor fiber) feeding at night, Knapdale Forest, Argyll, Scotland, UK, June.
90752765
90752765 Eurasian beaver (Castor fiber) feeding at night, Knapdale Forest, Argyll, Scotland, UK, June.© SCOTLAND: The Big Picture
Eurasian beaver (Castor fiber) feeding at night, Knapdale Forest, Argyll, Scotland, UK, June.
90752764
90752764 Eurasian beaver (Castor fiber) feeding at night, Knapdale Forest, Argyll, Scotland, UK, June.© SCOTLAND: The Big Picture
Trees and stumps with evidence of Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, site of Scottish Beaver Trial, Argyll, Scotland, UK, June.
90752761
90752761 Trees and stumps with evidence of Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, site of Scottish Beaver Trial, Argyll, Scotland, UK, June.© SCOTLAND: The Big Picture
Tree with signs of Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, site of Scottish Beaver Trial, Argyll, Scotland, UK, June.
90752760
90752760 Tree with signs of Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, site of Scottish Beaver Trial, Argyll, Scotland, UK, June.© SCOTLAND: The Big Picture
Tree felled by Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, site of Scottish Beaver Trial, Argyll, Scotland, UK, June.
90752759
90752759 Tree felled by Eurasian beaver (Castor fiber) feeding activity, Knapdale Forest, site of Scottish Beaver Trial, Argyll, Scotland, UK, June.© SCOTLAND: The Big Picture
Aerial view of European beaver (Castor fiber) habitat and  structures.  Beaver ponds, lodge, dams and felled trees  are visible , Spessart, Germany, April.
90737443
90737443 Aerial view of European beaver (Castor fiber) habitat and structures. Beaver ponds, lodge, dams and felled trees are visible , Spessart, Germany, April.© Ingo Arndt
Aerial view of European beaver (Castor fiber) habitat and  structures.  Beaver ponds, lodge, dams and felled trees  are visible , Spessart, Germany, April.
90737442
90737442 Aerial view of European beaver (Castor fiber) habitat and structures. Beaver ponds, lodge, dams and felled trees are visible , Spessart, Germany, April.© Ingo Arndt
Juvenile Eurasian beaver (Castor fiber) swimming in dammed stream in large enclosure, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, May.
90711546
90711546 Juvenile Eurasian beaver (Castor fiber) swimming in dammed stream in large enclosure, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, May.© Nick Upton
Downy birch tree (Betula pubescens) heavily gnawed by Eurasian beavers (Castor fiber) and close to being felled, in  large enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, May.
90711545
90711545 Downy birch tree (Betula pubescens) heavily gnawed by Eurasian beavers (Castor fiber) and close to being felled, in large enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, May.© Nick Upton
Adult male Eurasian beaver (Castor fiber) with identifying ear tag swimming at sunset among felled trees dammed stream within large enclosure, Devon Beaver Project, run by Devon Wildlife Trust, Devon,...
90711544
90711544 Adult male Eurasian beaver (Castor fiber) with identifying ear tag swimming at sunset among felled trees dammed stream within large enclosure, Devon Beaver Project, run by Devon Wildlife Trust, Devon,...© Nick Upton
Adult male Eurasian beaver (Castor fiber) with identifying ear tag swimming at sunset among felled trees dammed stream within large enclosure, Devon Beaver Project, run by Devon Wildlife Trust, Devon,...
90711543
90711543 Adult male Eurasian beaver (Castor fiber) with identifying ear tag swimming at sunset among felled trees dammed stream within large enclosure, Devon Beaver Project, run by Devon Wildlife Trust, Devon,...© Nick Upton
Eurasian beaver (Castor fiber) dam within woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, May.
90711541
90711541 Eurasian beaver (Castor fiber) dam within woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, May.© Nick Upton
Eurasian beaver (Castor fiber) using its front legs to lift and add mud and vegetation onto its dam, in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, May....
90711540
90711540 Eurasian beaver (Castor fiber) using its front legs to lift and add mud and vegetation onto its dam, in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, May....© Nick Upton
Willow tree (Salix sp.) felled and most of its bark stripped off by Eurasian beavers (Castor fiber) on the banks of the River Otter, Devon, UK, May.
90711539
90711539 Willow tree (Salix sp.) felled and most of its bark stripped off by Eurasian beavers (Castor fiber) on the banks of the River Otter, Devon, UK, May.© Nick Upton
Willow tree (Salix sp.) felled and most of its bark stripped off by Eurasian beavers (Castor fiber) on the banks of the River Otter, Devon, UK, May.
90711538
90711538 Willow tree (Salix sp.) felled and most of its bark stripped off by Eurasian beavers (Castor fiber) on the banks of the River Otter, Devon, UK, May.© Nick Upton
Eurasian beaver (Castor fiber) standing on its dam in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote camera trap.
90710810
90710810 Eurasian beaver (Castor fiber) standing on its dam in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote camera trap.© Nick Upton
Eurasian beaver (Castor fiber) standing on its dam in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote camera trap.
90710809
90710809 Eurasian beaver (Castor fiber) standing on its dam in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote camera trap.© Nick Upton
Tree felled by Eurasian beavers (Castor fiber) on the banks of the River Otter, Devon, UK, March.
90710805
90710805 Tree felled by Eurasian beavers (Castor fiber) on the banks of the River Otter, Devon, UK, March.© Nick Upton
Willow tree (Salix sp.) heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
90710804
90710804 Willow tree (Salix sp.) heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.© Nick Upton
Willow tree (Salix sp.) trunk heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
90710803
90710803 Willow tree (Salix sp.) trunk heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.© Nick Upton
Willow tree (Salix sp.) trunk heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
90710802
90710802 Willow tree (Salix sp.) trunk heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.© Nick Upton
Willow tree (Salix sp.) heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
90710800
90710800 Willow tree (Salix sp.) heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.© Nick Upton
Willow trees (Salix sp.) gnawed and felled by Eurasian beavers (Castor fiber), Wiltshire, UK, April.
90710799
90710799 Willow trees (Salix sp.) gnawed and felled by Eurasian beavers (Castor fiber), Wiltshire, UK, April.© Nick Upton
Young Willow tree (Salix sp.) felled by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.
90710798
90710798 Young Willow tree (Salix sp.) felled by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.© Nick Upton
Young Willow trees (Salix sp.) felled and gnawed by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.
90710797
90710797 Young Willow trees (Salix sp.) felled and gnawed by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.© Nick Upton
Young Willow tree (Salix sp.) felled by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.
90710796
90710796 Young Willow tree (Salix sp.) felled by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.© Nick Upton
Young Willow tree (Salix sp.) felled by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.
90710795
90710795 Young Willow tree (Salix sp.) felled by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.© Nick Upton
Canal made by Eurasian beavers (Castor fiber) in marshy woodland, within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April 2015.
90710788
90710788 Canal made by Eurasian beavers (Castor fiber) in marshy woodland, within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April 2015.© Nick Upton
Eurasian beaver (Castor fiber) gnawing the trunk of a Willow tree (Salix sp.) in woodland enclosure at night. Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote cam...
90710782
90710782 Eurasian beaver (Castor fiber) gnawing the trunk of a Willow tree (Salix sp.) in woodland enclosure at night. Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote cam...© Nick Upton
Eurasian beaver (Castor fiber) dam within a large woodland enclosure. Devon Beaver Project, Devon Wildlife Trust, Devon, UK, February 2015.
90710781
90710781 Eurasian beaver (Castor fiber) dam within a large woodland enclosure. Devon Beaver Project, Devon Wildlife Trust, Devon, UK, February 2015.© Nick Upton
Eurasian beaver (Castor fiber) arranging sticks on its dam in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote camera trap.
90710768
90710768 Eurasian beaver (Castor fiber) arranging sticks on its dam in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote camera trap.© Nick Upton
Eurasian beaver (Castor fiber) placing a branch stripped of bark on its dam in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote camer...
90710767
90710767 Eurasian beaver (Castor fiber) placing a branch stripped of bark on its dam in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote camer...© Nick Upton
Eurasian beaver (Castor fiber) placing a branch stripped of bark on its dam in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote camer...
90710766
90710766 Eurasian beaver (Castor fiber) placing a branch stripped of bark on its dam in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote camer...© Nick Upton
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