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90778292

90778292 Leopard resting with cub (Panthera pardus) Masai Mara NR, Kenya© Anup Shah

Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years crossing a river, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic

90776574

90776574 Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years crossing a river, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic© Anup Shah

90704178

90704178 Female Cheetah scans horizon for prey {Acinonyx jubatus} Masai Mara, Kenya© Anup Shah

Bornean Orangutan (Pongo pygmaeus wurmbii) female 'Princess' walking along a trail carrying her daughter 'Putri' aged 2 years on her back. Camp Leakey, Tanjung Puting National Park, Central Kalimantan...

90763559

90763559 Bornean Orangutan (Pongo pygmaeus wurmbii) female 'Princess' walking along a trail carrying her daughter 'Putri' aged 2 years on her back. Camp Leakey, Tanjung Puting National Park, Central Kalimantan...© Anup Shah

Bornean Orangutan (Pongo pygmaeus wurmbii) female 'Tata' and her unnamed baby aged 2-3 months portrait. Camp Leakey, Tanjung Puting National Park, Central Kalimantan, Borneo, Indonesia. June 2010. Reh...

90763556

90763556 Bornean Orangutan (Pongo pygmaeus wurmbii) female 'Tata' and her unnamed baby aged 2-3 months portrait. Camp Leakey, Tanjung Puting National Park, Central Kalimantan, Borneo, Indonesia. June 2010. Reh...© Anup Shah

90731657

90731657 Lioness (Panthera leo) rejecting a male's advances Maasai Mara National Reserve, Kenya.© Anup Shah

Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years and juvenile male 'Mobangi' aged 5 years drinking from a dirty pool of water in Bai Hokou, Dzanga Sangha Specia...

90730946

90730946 Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years and juvenile male 'Mobangi' aged 5 years drinking from a dirty pool of water in Bai Hokou, Dzanga Sangha Specia...© Anup Shah

90729650

90729650 Sumatran Orang utan male in canopy calling out (Pongo abelii) Gunung Leuser NP, Sumatra, Indonesia© Anup Shah

Spotted hyena (Crocuta crocuta) aged about 2 years watching with curiosity with the rest of the clan in the background. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

90726057

90726057 Spotted hyena (Crocuta crocuta) aged about 2 years watching with curiosity with the rest of the clan in the background. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.© Anup Shah

White-backed vultures (Gyps africanus) waiting for Spotted hyenas (Crocuta crocuta) to finish feeding on a kill. Maasai Mara National Reserve, Kenya.

90726056

90726056 White-backed vultures (Gyps africanus) waiting for Spotted hyenas (Crocuta crocuta) to finish feeding on a kill. Maasai Mara National Reserve, Kenya.© Anup Shah

90726055

90726055 Spotted hyena (Crocuta crocuta) approaching with curiosity. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.© Anup Shah

90726054

90726054 Spotted hyena (Crocuta crocuta) walking and smelling the ground. Maasai Mara National Reserve, Kenya.© Anup Shah

Tana mangabey (Cercocebus galeritus) mother grooms her baby of between -3 months old watched by a juvenile. Tana River Forest, South eastern Kenya.

90722562

90722562 Tana mangabey (Cercocebus galeritus) mother grooms her baby of between -3 months old watched by a juvenile. Tana River Forest, South eastern Kenya.© Anup Shah

90722561

90722561 Eastern White-bearded Wildebeest (Connochaetes taurinus) herd crossing the Mara River. Maasai Mara National Reserve, Kenya.© Anup Shah

African elephant (Loxodonta africana) herd at waterhole with calves seen between adults legs. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

90720592

90720592 African elephant (Loxodonta africana) herd at waterhole with calves seen between adults legs. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.© Anup Shah

Eastern white-bearded wildebeest (Connochaetes taurinus) herd running. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

90720591

90720591 Eastern white-bearded wildebeest (Connochaetes taurinus) herd running. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.© Anup Shah

African elephants (Loxodonta africana) wallowing in a waterhole. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

90720589

90720589 African elephants (Loxodonta africana) wallowing in a waterhole. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.© Anup Shah

90720588

90720588 African elephants (Loxodonta africana) dust bathing. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.© Anup Shah

Black-backed jackals (Canis mesomelas) feeding on a wildebeest carcass. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

90720587

90720587 Black-backed jackals (Canis mesomelas) feeding on a wildebeest carcass. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.© Anup Shah

90720586

90720586 Lion (Panthera leo) old male with one eye standing next to a kill. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.© Anup Shah

Sumatran orangutan (Pongo abelii) female 'Juni' aged 12 years swinging from a liana. Gunung Leuser National Park, Sumatra, Indonesia. Rehabilitated and released (or descended from those which...

90720469

90720469 Sumatran orangutan (Pongo abelii) female 'Juni' aged 12 years swinging from a liana. Gunung Leuser National Park, Sumatra, Indonesia. Rehabilitated and released (or descended from those which...© Anup Shah

90720467

90720467 Lion-tailed macaque (Macaca silenus) male yawning portrait. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India.© Anup Shah

Lion-tailed macaque (Macaca silenus) female sitting with her baby aged 12-18 months. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India.

90720466

90720466 Lion-tailed macaque (Macaca silenus) female sitting with her baby aged 12-18 months. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India.© Anup Shah

Lion-tailed macaque (Macaca silenus) female sitting with her baby aged less than 1 month. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India.

90720465

90720465 Lion-tailed macaque (Macaca silenus) female sitting with her baby aged less than 1 month. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India.© Anup Shah

90720464

90720464 Lion-tailed macaque (Macaca silenus) male yawning. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India.© Anup Shah

90720463

90720463 Lion-tailed macaque (Macaca silenus) baby aged less than 1 month playing in a tree. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India.© Anup Shah

90720462

90720462 Lion-tailed macaques (Macaca silenus) play fighting. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India.© Anup Shah

90720109

90720109 Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer). Masai Mara National Reserve, Kenya, July© Anup Shah,Anup Shah

90720108

90720108 Lioness (Panthera leo) with cub aged 2-3 months portrait. Masai Mara National Reserve, Kenya, August© Anup Shah

90720107

90720107 Masai giraffe (Giraffa camelopardalis tippelskirchi)male in front of woodland, Masai Mara National Reserve, Kenya, July© Anup Shah

90720106

90720106 Spotted hyena (Crocuta crocuta) preparing to carry a pup aged 2-3 months. Masai Mara National Reserve, Kenya, July© Anup Shah

Olive baboon (Papio cynocephalus anubis) female with infant on back, carrying away a gazelle carcass having taken it from a male. Masai Mara National Reserve, Kenya, July

90720105

90720105 Olive baboon (Papio cynocephalus anubis) female with infant on back, carrying away a gazelle carcass having taken it from a male. Masai Mara National Reserve, Kenya, July© Anup Shah,Anup Shah

90720104

90720104 Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer). Masai Mara National Reserve, Kenya, July© Anup Shah,Anup Shah

90720103

90720103 Lion (Panthera leo) male nuzzling lioness, Masai Mara National Reserve, Kenya, July© Anup Shah

90720102

90720102 Eastern White-bearded Wildebeest (Connochaetes taurinus) herd on the move Masai Mara National Reserve, Kenya, July© Anup Shah,Anup Shah

Eastern White-bearded Wildebeest (Connochaetes taurinus) herd crossing the Mara River and clambering up steep bank. Masai Mara National Reserve, Kenya, July

90720100

90720100 Eastern White-bearded Wildebeest (Connochaetes taurinus) herd crossing the Mara River and clambering up steep bank. Masai Mara National Reserve, Kenya, July© Anup Shah

90720091

90720091 Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer) Masai Mara National Reserve, Kenya© Anup Shah,Anup Shah

Cheetah (Acinonyx jubatus) cub aged about one year running. Maasai Mara National Reserve, Kenya.

90704615

Japanese Macaque (Macaca fuscata) baby sitting on the edge of thermal hotspring pool. Jigokudani Yean-Koen National Park, Japan, February.

90704483

90704483 Japanese Macaque (Macaca fuscata) baby sitting on the edge of thermal hotspring pool. Jigokudani Yean-Koen National Park, Japan, February.© Anup Shah

90699511

90699511 Topi (Damaliscus lunatus jimela) running. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.© Anup Shah

90699510

90699510 African elephant (Loxodonta africana) close-up. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.© Anup Shah

90699506

90699506 Spotted hyena (Crocuta crocuta) approaching with curiosity. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.© Anup Shah

90699504

90699504 Olive baboon (Papio anubis) sitting portrait. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.© Anup Shah

Celebes / Black crested macaque (Macaca nigra) two juveniles approaching with curiosity, one grimacing Tangkoko National Park, Sulawesi, Indonesia.

90698798

90698798 Celebes / Black crested macaque (Macaca nigra) two juveniles approaching with curiosity, one grimacing Tangkoko National Park, Sulawesi, Indonesia.© Anup Shah

90698797

90698797 Celebes / Black crested macaque (Macaca nigra) group up close, watching with curiosity, Tangkoko National Park, Sulawesi, Indonesia.© Anup Shah

Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years and juvenile male 'Mobangi' aged 5 years approaching with curiosity, Bai Hokou, Dzanga Sangha Special Dense For...

90698580

90698580 Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years and juvenile male 'Mobangi' aged 5 years approaching with curiosity, Bai Hokou, Dzanga Sangha Special Dense For...© Anup Shah / naturepl

Young African elephant holding plant with trunk, Masai Mara, Kenya - Anup Shah/ npl

90141096

Chimpanzees gather round alpha male 'Frodo' in tree feeding on dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002 - Anup Shah/ npl

90141012

90141012 Chimpanzees gather round alpha male 'Frodo' in tree feeding on dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002© Anup Shah / NPL

Hippopotamus (Hippopotamus amphibius) walking through grass. Maasai Mara National Reserve, Kenya.

90872291

90872165

90872165 Lion-tailed macaque (Macaca silenus) baby aged 9-12 months portrait. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India.©

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